Tim,
I can't really answer the question, but i think i ran into something like this in the past.
I expect you went to "Implementation Manager" in the original Test Tenant, and did "Download A Copy"
You then logged into the new Test Tenant and did "Upload" of the copy that you had downloaded from the original Test Tenant.
What is the Status of your Project in the new Test Tenant? Where you see Solution_Name (SolutionStatus) within the "Project Explorer", does it actually still say "In Development" in the new Test Tenant? I seem to recall when i did this process, it changed the status to something that wasn't "In Development"